International Arctic Research Center, University of Alaska Fairbanks, 930 Koyukuk Dr., Fairbanks, AK 99775, United States
School of Life and Environmental Sciences, Osaka Prefecture University, 1-1 Gakuen- cho, Naka-ku, Sakai, Osaka, 599-8531, Japan ; International Arctic Research Center, University of Alaska Fairbanks, 930 Koyukuk Dr., Fairbanks, AK 99775, United States
School of Life and Environmental Sciences, Osaka Prefecture University, 1-1 Gakuen- cho, Naka-ku, Sakai, Osaka, 599-8531, Japan ; International Arctic Research Center, University of Alaska Fairbanks, 930 Koyukuk Dr., Fairbanks, AK 99775, United States