University of California Berkeley, Department of Integrative Biology, Valley Life Sciences Building, Berkeley, CA 94720-3140 United States
Northern Arizona University, School of Forestry, Flagstaff, AZ 86011-5018 United States
Northern Arizona Univeristy, Department of Biological Sciences and Merriam Powell Center for Environmental Research, Flagstaff, AZ 86011-5018 United States
Northern Arizona Univeristy, Department of Biological Sciences and Merriam Powell Center for Environmental Research, Flagstaff, AZ 86011-5018 United States